155-10-01: Roxburghii, are you in there??

For years there has been much discussion about the pedigree of a number of roses that list the species R. roxburghii as a parent. The Tantau Floribunda 'Floradora' (1944, Germany), for example, lists 'Baby Château' as the seed parent and R. roxburghii as the pollen parent.

Now, at a glance, there isn't the slightest hint of the species parent in the look of 'Floradora', which has led many to doubt its listed parentage. I myself doubted Tantau's pedigree for the longest time, at least unbtil I had some experience in using certain species and near-species hybrids in breeding, especially when they were employed as pollen parents. Let me elaborate.

I frequently use certain seed parents in making crosses that might be impossible on many other parents. After a while, you start to recognize which of your seed parents are likely to be "door openers" when planning iffy crosses. Introducing certain species can be especially difficult; sometimes the chromosome counts don't match, or the two varieties are simply too distant from one another in the family. I believe there are other unknown factors that play a role in fertility. Whatever the case, I found that even when using very willing seed parents, certain pollens would result in seed that, once germinated, turned out to be the result of apomixis*.

Apomixis is what happens when pollen initiates the fertilization response in a plant, but once the pollen tubes grow into the ovary, the genes prove unusable. So, in an effort to salvage the seed making process, the plant instead duplicates its own genes, in effect cloning itself. In roses, I suspect there is some recombinant action occurring as well, since some of the individuals arising from such crosses are not identical to the parent plant, but appear to be highly similar. In some cases undoubtedly some selfing may also occur, perhaps the result of insect activity, or simply incomplete emasculation.

The seedling you see pictured here was an experiment designed to see what happens when a (supposed) second generation R. roxburghii hybrid (in this instance, Moore's 0-47-19 was used) was crossed once again with the species R. roxburghii. And so, what we have here is:

[R. wichurana X ('Baby Château' X R. roxburghii)] X R. roxburghii normalis. Clearly, assuming the Tantau parentage is correct, we have a lot of species genes in this plant. Now, when I made this cross I expected one of three possible scenarios: total rejection of the pollen, apomixis, or self-pollination by insect vectors. As it happens, I got only a few hips from the 40 plus pollinations, leading me to believe they were likely the result if insect fiddling. From the approximately 20 seeds I sowed I got two seedlings. The one illustrated here is the healthier of the two; the other is far less vigorous, but it is trying to build up steam. (Maybe our cold, prolonged spring isn't to its liking; I know its not making me feel great!)

Until this individual reached a certain size I was unwilling to make many assumptions about its pedigree. But now, I look at it and I come to the conclusion that this is actually, for real, a R. roxburghii hybrid. Look closely: the most recent leaves are now composed of nine leaflets, surely a sign that roxburghii is influencing its development. I don't expect to see flowers this year, which is fine. Right now I am just fascinated to see its vegetative development. I doubt this is any kind of proof of the presence of R. roxburghii in its ancestor, 'Floradora', but it does show that when using the species, the resulting seedlings can show clear evidence of roxburghii traits. Fascinating.

*For more on apomixis, see the Wikipedia article here.

170-09-02

17-09-02 = R. arkansana X 'Carlin's Rhythm'.

I'm determined to obtain a seedling from the delicate-looking-but-unbelievably-tough R. arkansana, a North American native. The species has richly scented single candy pink blooms, and certain select clones sometimes have subtle stripes or streaks of darker color on the petals. Mine doesn't, but mine does do something I hope to capitalize on; it blooms at least three times in the growing season: late May, again in July or early Augustm then again in late September or October.

My specimen of R. arkansana is a reluctant seed setter, rarely accepting pollen from anything but itself. However, I did manage to get two seedlings from a cross using Kim Rupert's beautiful 'Carlin's Rhythm'. This is the second of the two (the first was paler and not as nice) and displays large-ish blooms (about 2.5") in a deeper than average rose pink. The scent is rich and pure "old rose". Now, the trick is to try it in breeding to see if I can make the next step. It will have one copy of the modern hybrid remontancy gene, and at least once copy of the gene that makes my arkansana repeat, which may or may not be distinct from other forms of remontancy. As both parents are (at least in principle) tetraploids, I would expect this seedling to be a tetraploid also.

220-09-02

Apparently the striping is a fixed trait; the next bloom to open is identical. Excellent.

220-09-02

220-09-02: "A supposed tetraploid form of R. rugosa" X Basye's Blueberry.

The seed parent is a rose shared with me by Joan Monteith about a decade ago. Joan did an experiment in which she treated a select form of R. rugosa with the chemical Colchicine with the hopes of doubling the plant's gene count from 2X to 4X, making it more compatible with most modern tetraploid hybrids. It has not been determined with any certainty that this Rugosa is in fact a Colchicine-induced tetraploid; measurements of pollen diameter has been inconclusive. However, I have pursued breeding concepts that presume it a tetraploid and I will simply evaluate the results as I proceed.

Case in point; today's seedling. 220-09-02 is the first seedling from this "presumed tetra Rugosa" X 'Basye's Blueberry' cross. Of the dozen or so seedlings I got from the cross, remarkably not a one has the classic Rugosa foliage. In fact, most all lean heavily towards the look of the Basye parent, which comes as a huge surprise. (Most R. rugosa hybrids in the first generation show strong Rugosa influence, often obliterating all of the other parent's qualities)

The flower is small, at just under 2", quite heavily Clove-scented, thick of petal and most remarkably, each petal has a strong white streak down the middle. Don't ask me where this trait came from. I do know that Ralph Moore once showed me a Rugosa hybrid with about 15 petals that showed the same petal streak, and it was striking. Pollen has been gathered from this seedling and will be placed on a number of tetraploid seed parents I have assigned for the purpose of testing pollen fertility. In the meantime, I look forward to evaluating this plant for sturdiness and disease resistance. I am hopeful, given its pedigree, that Blackspot resistance might be superb.

165-09-03

165-09-03: another approach to working with R. foliolosa. Click on the photo to view a larger version.

A few days ago I mentioned 44-09-13, one of my Foliolosa/Therese Bugnet hybrids. In the case of 44-09-13, the seed parent was a second generation R. foliolosa hybrid, whereas with today's seedling, 165-09-03, R. foliolosa itself plays that role. Both crosses utilize Therese Bugnet as the pollen donor. The idea was simply to make some Therese Bugnet crosses using any/all diploids I had on hand and which I knew to be decent seed bearers. The R. foliolosa approach was, to me, particularly appealing since it was such a healthy, Winter hardy individual. R. foliolosa also imparts on its progeny Fall foliage coloring to make the most dramatic of Maples and Sumacs envious; fiery oranges, yellows overlaid with flame red. I have come to think that modern roses ought to have some degree of "four season appeal".

I consider the seedling pictured here to be pretty much intermediate between its parents, although the flower itself leans more towards Therese Bugnet in size and petal count. The bloom has a moderately intense Rugosa scent; a welcome trait, to be sure. (The weather has been abysmally cold this Spring, so maybe when/if it warms up, the fragrance will be more intense.) The foliage could easily be said to resemble either parent, since both have fern-like feathery foliage that is narrow of leaflet and pleasantly matte in texture.

Pollen has been collected from both this seedling and its relative, 44-09-13, to be tested for fertility this year. I will try to limit myself to working it on other diploids and maybe a triploid or two, but I might just dust a few blooms of Midnight Blue while I'm at it. I won't let the creation of triploids (and their potential for sterility) stop me from making a cross once I have a few foundation plants established in the confines of a given ploidy. As Ralph so often said, "The rose will find a way."

Tilda Swinton says birth is 'violent'

Tilda Swinton claims Hollywood glamorises childbirth, when in fact it is a ''violent'' experience for women.

Tilda Swinton thinks childbirth is "violent"

The 50-year-old actress - who has 13-year-old twins Honor and Xavier with partner John Byrne - believes Hollywood glamorises birth, which is actually a "murderous" experience.

She said: "In movies, and particularly in television films, when people have babies they are sitting in a hospital room and there are flowers everywhere.

"They are made up, magically, and they have a baby in their arms and it's all really lovely. They start getting pink baby clothes. It's a truly, truly murderous business. It's violent."

The Oscar-winning actress - who portrays a mother who fails to bond with her son in forthcoming movie 'We Need to Talk About Kevin' - thinks women who are unable to "embrace" the nature of child birth may find themselves "up a gum tree" because of how hard it is.

Speaking at the Cannes Film Festival, she said: "If one doesn't embrace that [violence], if one can't embrace it - and it's really tough to do that - then you're up a gum tree because it means you're going to be cutting off a whole part of yourself."

Tags: Tilda Swinton, Violent, Glamorises, Television films, Mouderous, Embrace

44-09-13

44-09-13 = (R. foliolosa X 'Little Chief') X 'Therese Bugnet'

This is the first of this group of seedlings to bloom. Hardly an exciting flower, really, but not a surprising result, really, considering how significant a role R. rugosa plays in its pedigree. The fragrance, in fact, is pure Rugosa: richly Clove-like. The foliage is pure 'Therese Bugnet'; ferny, with narrow leaflets, and that odd matte texture. The foliage has been impermeable to the standard diseases so far, but testing in earnest for Blackspot won't happen till the selections are planted out in the open garden.

For now, I am collecting pollen from some of these selections to test as breeders. I don't consider these individuals as "finished" works, but as potential stepping stones towards improved cold climate-friendly plants.

You can view info about the seed parent here.